Veiled Chameleon Food Pyramid
The pyramid is a guide to feed your veiled chameleon. You do not have to follow the schedule exactly (see special considerations). It is based off of research on the macronutrient and calcium requirements of various types of reptiles outlined in the Merck Veterinary Manual as well as individual scientific studies about wild veiled chameleon diets. This pyramid is also based off of what experienced veiled chameleon breeders have found to work for their animals.
The veiled chameleon food pyramid is designed to be a user-friendly guide. The diagram in the insect section shows how feeder insects should be properly gut loaded for a veiled chameleon. These amounts are proportioned by weight. The key on the right side describes how many times to feed this food group per a week. When the pyramid level is broken in half, you can choose either food group. The larger the section, the more important it is. |
Introduction: |
This guide is made to combat the incomplete knowledge of veiled chameleon nutrition. Many new owners are confused about what to feed their pets while experienced owners are looking for the best diet possible.
There is no such thing as a complete guide to veiled chameleon nutrition because there is no realistic way for the in-home keeper to formulate a total diet. It is conventional wisdom that variety is the key to combat an incomplete diet, but that isn’t always the case. For example, almost all the foods a veiled chameleon is fed will be deficient in calcium and iodine. This is simply a product of commercial agriculture raising plants and insects so that they grow as cost effectively as possible. This guide will get you as close as possible to a complete diet without expensive scientific testing. The most important thing in any animal’s diet is feeding appropriate foods in appropriate ratios in order to get the correct balance of protein fat and carbohydrates. It will also help you understand other nutrients how to get them too. |
Key Food Groups:
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Insects:
Insects are the protein source for veiled chameleons. Most commercially raised insects are deficient in several nutrients, including calcium, thiamin (vitamin B1), vitamin A and vitamin E. It is necessary to feed the insects a high calcium and vitamin A diet before feeding to your veiled chameleon. Gut loading is much more effective than dusting insects. The insects fed should also be low in fat content. Chameleons of all species have a preference for a variety of insects. Studies have been conducted on the common chameleon and the jackson's chameleon showing that they eat a wide variation of arthropods. Of these arthropods most are insects (Pleguezuelos et al. 1999 and Kraus and Preston 2012). Eason (1990) showed that the senegalese chameleon preferred insects that it had not consumed recently over insects that it had recently eaten. This adaptation is certainly to ensure a diet that has a large variation and should be mimicked as much as possible in captivity. Ideal insects include crickets, roaches, hornworms and soldier fly larva (reptiworms). Insects that should be fed more sparingly due to their fat content include mealworms , superworms and waxworms. Check out the nutritional analysis of some common feeder insects. Insects' Diet- Gut Loading: Gut loading insects is critical for all chameleons because so much, if not all, of their diet comes from insects. The food for gut loading should be 20% calcium, 50% greens, 10% vitamin/mineral supplement and 20% vegetables, by weight. This diet should be fed to the insects for at least 48 hours. This allows the body to absorb nutrients such as vitamin A. Calcium is not well consumed by insects, so in addition to feeding this gut load diet, insects should be dusted with a calcium powder. -Calcium (20% of gut loading diet): A calcium supplement should be purchased separately from a vitamin and mineral supplement. Almost all feeder insects are deficient in calcium and calcium is required more than any other mineral in reptiles. The calcium supplement fed to insects should include vitamin D3 to ensure that the insects can process the calcium. The calcium recommendations for gut loading are based on the Finke (2003) study that recommends 3-9% elemental calcium in a gut loading diet, depending on the insect species. Calcium carbonate (the most common calcium supplement) is 40% elemental calcium, thus 20% calcium supplement is needed in the diet. This recommendation is also backed up by Finke et al. (2005), which showed that 15% calcium carbonate supplementation to the gut loading diet increased the calcium content of small crickets from 0.2% to 1.1% calcium. Considering that the 20% calcium supplement amount is by weight, it actually won't appear to be as much calcium as you would expect. It is highly recommended you weigh out the ingredients in your gut loading diet. Due to palatability issues with simply adding a calcium powder to a gut load diet of fresh vegetables, insects should also be dusted with a calcium powder before feeding. It is unlikely that insects will consume enough calcium in a home-made gut load diet. For the chameleons be sure that you have proper UVB lighting or dietary vitamin D supplementation. Dietary vitamin D3 supplementation can be dusted directly on insects and other food items. Without UVB or dietary vitamin D, veiled chameleons cannot process the calcium they are eating. UVB lighting is by far the better option because it is difficult to overdose, but Hoby et al. (2010) proved that either vitamin D source is effective in veiled chameleons. Significant research has been conducted into the calcium metabolism of the veiled chameleon. In growing chameleons that were supplemented with calcium and vitamin A, metabolic bone disease did not develop. This finding was independent of if they were being supplemented with dietary vitamin D or UVB lighting (Hoby et al. 2010). This indicates that retinol supplementation may be more important that the source of vitamin D in veiled chameleons. These claims are further backed up by Haxhiu et al. (2014) studies on expression of the vitamin D dependent calcium transport system in the duodenum of the veiled chameleon. Basically, higher expression means more dietary calcium will be absorbed. Supplementing UVB without calcium and without vitamin A supplementation yields the same low expression levels as no UVB supplementation at all. Significantly higher levels were expressed when calcium, vitamin A, vitamin D and UVB were all supplemented together. -Vitamin/Mineral Supplement (10% of gut loading diet): A vitamin and mineral supplement is essential when feeding insects to ensure no nutrients are missing, however there is quite a variety of supplements on the market. When looking for a supplement for a healthy gut loading plan, as outlined here, there are a few key nutrients one should consider. Most insects are low in magnesium, iron, iodine and vitamin E. Make sure your supplement provides these nutrients as well as ample amounts of B vitamins. Often iodide is provided using kelp as an ingredient. Vitamin A is also a nutrient commonly lacking from commercially raised insects. A vitamin and mineral supplement can be used to help increase the vitamin A content of a gut loading diet, but if fed the correct vegetables in the gut loading diet, this is not required. Greens: Greens are a powerful source of nutrition for veiled chameleons. They also provide moisture, bulk and vitamins to a gut loading diet. Greens generally can be divided into three categories, staple, occasional rare and never foods. To make a green an every day green it must be rich in nutrients like calcium, low in phosphorus and low in anti-nutrients like oxalates and goitrogens. Occasional greens are rich in nutrients, but contain too high a level of oxalates or goitrogens to be feed every day. Rare greens have very high oxalates and goitrogens. They should be avoided. Never greens have high phosphorus, low calcium and sometimes have high oxalates. Staple greens are escarole, endive, alfalfa and dandelion greens. Occasional greens are bok choy, collard greens, mustard greens, turnip greens, watercress Rare greens are kale, dark lettuces, parsley, swiss chard, spinach Never greens are iceburg lettuce, cabbage Vegetables: This is the part of a veiled chameleon’s diet that will bring in a diversity of nutrients and flavors. Vegetables are also critically important for gut loading as a source of beta-carotene, which is processed by the insects into retinol, a form of vitamin A that veiled chameleons can use. Good vegetables are often bright in color and not too high in oxalates and phosphorus. Sometimes the healthiest foods are also the strangest, such as prickly pear leafs and hibiscus flowers. Some of the best vegetables include acorn squash, butternut squash, bell peppers, prickly pear leafs, hibiscus, common button mushroom, okra, and pumpkin. Other good vegetables include asparagus, broccoli, brussel sprouts, carrot, green beans, snap peas, tomato, yellow squash and cucumber. Check out the nutritional analysis of vegetables. Fruit: Fruit can be fed to veiled chameleons but does not appear on the pyramid because not only is fruit a rare occurrence in their wild diet, almost all fruit you can buy in the store is heavily domesticated and has little relation to the nutritional content of its wild relatives. Fruit should be used as a flavor enhancer and very rare treat. Some healthier fruits include berries, cantelope, dates, figs, guava, kiwi, mango, oranges, papaya and pineapple. Sources: Eason, Perri K. The Effect of Recent Diet on Prey Choice in Senegalese Chameleons (Chamaeleo senegalensis). Journal of Herpetology. 24(4): 383-387 (1990). Finke, Mark D. Gut Loading to Enhance the Nutrient Content of Insets as Food for Reptiles: A Mathematical Approach. Zoo Biology. 22(2):147-162 (2003). Finke, Mark D., Shari U, Dunham and Christabel A. Kwabi, Evaluation of Four Dry Commercial Gut Loading Products for Improving the Calcium Content of Crickets, Acheta domestics. Journal of Herpetological Medicine and Surgery. 15(1) 7-12 (2005). Haxhiu, D., S. Hoby, C. Wenker, A. Boos, M. P. Kowalewski, F. Lewis and A. Liesegang. Influence of Feeding and UVB Exposition on the Absorption Mechanisms of Calcium in the Gastrointestinal Tract of Veiled Chameleons (Chamaeleo calyptratus). Journal of Animal Physiology and Animal Nutrition. 98: 1021-1030 (2014). Hoby, Stefan, Christian Wenker, Nadia Robert, Thomas Jermann, Sonja Hartnack, Helmut Segner, Claude-P Aebischer and Annette Liesegang. Nutritional Metabolic Bone Disease in Juvenile Veiled Chameleons (Chamaeleo calyptratus) and Its Prevention. American Society for Nutrition. 140(11): 1923-1931 (2010). Kraus, Fred and David Preston. Diet of the Invasive Lizard Chamaeleo jacksonii (Squamata: Chamaeleonidae) at a Wet-Forest Site in Hawaii. Pacific Science. 66(3): 397-404 (2012). Pleguezuelos, Juan M., Juan C. Poveda, Raquel Monterrubio and Diego Ontiveros. Feeding Habits of the Common Chameleon, Chamaeleo Chamaeleon (Linnaeus 1758) in the Southeastern Iberian Peninsula. Israel Journal of Zoology. 45(2): 267-276 (1998). |
Special Considerations:
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These dietary guidelines are designed for an average, healthy, veiled chameleon. As with anything, if your reptiles are ill you need to consult a veterinarian for advice. That being said, there are some normal dietary considerations that apply to many veiled chameleons. If your veiled chameleon is sick, please check out our diagnostics page.
Obesity: Many captive animals of all kinds suffer from obesity. The best thing you can do for an obese veiled chameleon is feed only lean insect species and stop feeding fruit. Obesity is deadly. It can cause liver failure and many other issues. Supplementing with a very small amount of omega 3 fish oil is also wise to prevent liver disease in obese animals. Juveniles: Young veiled chameleons are growing fast and need more protein and calcium than older veiled chameleons. It is wise to increase the amount of insects offered to them. Fatty insects are also a good source of energy for young animals. Make sure all insects fed are gut loaded with calcium and vitamin A. Anorexia: Not eating is a common occurrence with many pet reptiles. Usually the issue is with the temperature of the enclosure, or another husbandry problem. Ensure that your tank is up to standards and visit a veterinarian. In the mean time, remember that eating any food is better than eating no food. Even if all your veiled chameleon will eat is banana baby food this is better than not eating anything at all. Work on getting them to eat healthy foods, but again, unhealthy food is better than no food. For tips and tricks to get your animal to eat see our "appetite stimulation" page. |
Key Nutrients:
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There are many key nutrients in a veiled chameleon diet. Calcium is a particularly important nutrient that many beginner keepers do not offer enough of. It is imperative to supplement your veiled chameleon with calcium and provide proper UVB lighting.
The key nutrients are listed in alphabetical order. Anthocyanin Definition-natural plant flavonoid pigments that appear blue or purple Function-a weak antioxidant that enhances color Foods-acai, blood orange, blackberry, raspberry, blueberry, purple grapes Beta-carotene Definition-a red-orange pigment in plants that can be converted by some animals to vitamin A Function-A vitamin A supplement for omnivores and herbivores that cannot be overdosed as well as an antioxidant that enhances color Foods-carrot, pumpkin, sweet potato, spinach Calcium Definition-an abundant chemical element that essential for life Function-building blocks for bones and teeth and an essential element for cell physiology Foods-calcium carbonate supplement, prickly pear leafs, collard greens, oranges, soldier fly larva, mustard greens, bell peppers Carbohydrates Definition-a biological molecule made of chemically linked sugars. Carbohydrates include, sugars, starch and fiber Function-simple energy source for animals Foods-almost all foods, especially plants Copper Definition-a chemical element that is an essential mineral Function-Copper has many functions, including collagen formation, enzyme cofactors, incorporating iron into red blood cells and generation of energy from carbohydrates. It is usually abundant in foods for veiled chameleons and inhibits zinc uptake, so copper intake needs to be limited. Foods-spinach, turnip greens. asparagus, beans, nuts, kale, mushroom, pumpkin Fats (lipids) Definition-a biological molecule that is found in plants and animals Function- a dense energy source and an energy storage method Foods-waxworms, superworms, mealworms, nuts, seeds Fiber - Click Here for More Information Definition-a type of complex carbohydrate in plants that cannot be digested Function-Fibers add bulk to the stool and give food for gut bacteria Foods- all vegetables and fruit Goitrogens Definition-substances that disrupt the production of thyroid hormones Function-These anti-nutrients should be avoided in excess in order to maintain thyroid health of veiled chameleons. They are less harmful when proper iodine supplementation is provided. Foods-cassava, soy, peanuts, strawberries, spinach, bok choy, broccoli, cabbage, kale, turnip Iodine Definition-a chemical element abundant in the ocean that is an essential mineral Function-key component of hormones made by the thyroid Foods-kelp, ocean fish, shrimp, spirulina, scallops, standard table salt Macronutrient Definition- A category of nutrients that makes up the bulk of foods. It includes protein, fat(lipids), carbohydrates and water. Function-each macronutrient has a different function, but protein, fat(lipids) and carbohydrates can all provide energy. The ratios of these nutrients in a diet is extremely important. Foods-all foods Minerals (dietary) Definition-a chemical element that is required by living organisms other than carbon, hydrogen, nitrogen and oxygen Function-builds bones, tissue, catalyzes biological reactions, transports oxygen and many more Foods-all foods have some mineral content Oxalates Definition-a specific organic acid found in plants and animals Function-binds with calcium to form kidney stones and prevent calcium from being used in the body Foods-soy, nuts, beans, spinach, swiss chard, beets, collard greens, okra, blueberries, grapes, raspberries Phytonutrients Definition-chemicals naturally produced by plants that are not of conventional nutritional value Function-functions vary but they are usually antioxidants, anti-inflammatory and promote liver health Foods-plants, especially ones rich in color Protein - Click Here for More Information Definition-a biological molecule made of chemically linked amino acids Function-maintains the structure of the body and excess can be used as an energy source Foods-all foods have some protein, with insects being especially high as well as a decent amount of protein in nuts and seeds Phosphorus Definition- an abundant chemical element that essential for life Function-enables cell function and builds bones, but is easily consumed in excess in a veiled chameleon’s diet. Excess consumption impairs calcium absorption and can cause muscle disorders. Foods-asparagus, beans, soy, seeds, cricket, mealworms, mushroom Vitamins Definition-A vital organic compound that an organism cannot synthesize on its own. They can be divided into fat soluble and water soluble. Function-each vitamin has its own function in the body Foods-all foods Vitamin A Definition-a group of related fat-soluble nutrients (carotenoids and retinoids) that are essential for the body. Function- Carotenoids come from plants and cannot be overdosed, so they are the safest form of vitamin A, however it is not known if veiled chameleon’s can convert carotenoids to retinoids. Vitamin A is needed for immune responses, night vision and many other functions in the body. Foods-egg yolk, turkey liver Vitamin D Definition-a fat soluble vitamin derived from UVB light or dietary sources Function- needed to absorb calcium and other minerals from food Foods-UVB lighting should be provided for veiled chameleons so that they can synthesize vitamin D. Studies have shown they can also absorb dietary vitamin D, which is only readily available in commercial supplements. Vitamin E Definition-a fat soluble vitamin that acts as a powerful antioxidant Function-protects against free radical damage, ageing and heart disease. It is especially crucial in a high fat diet. Most foods high in vitamin E are also high in fat, so a balance is difficult to achieve. Foods-sunflower seeds, shrimp, greens. almonds, broccoli |
Science Behind the Pyramid:
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The proportions of the pyramid are based on macronutrient balancing. According to the Merck Veterinary Manual, omnivorous reptiles need a diet of 20-25% protein, 3-6% fat and 20-35% fiber on a dry matter basis. According to this manual carnivorous reptiles need 30-50% protein. Considering veiled chameleons are almost carnivorous the target for protein is at least 30%. To achieve these macronutrient ratios the pyramid recommends how often to feed a portion of insects, greens, fruits or vegetables. There is a total of 10 portions of these food groups fed each week.
.75 greens portions/6 total portions=12.5% greens .25 vegetable portions/6 total portions=4.2% vegetables 5 insect portions/6 total portions=83.3% insects The most important macronutrient is protein. The diet aims to achieve a >30% dry matter protein content. Greens, vegetables and insects are typically 20%, 10% and 50% protein, respectively. Multiplying the % of protein by the proportion of portions gives us: .125x20% + .042x10% + .833x50%= 44.6% protein Therefor this food pyramid provides an adequate amount of protein. Using this same method fat and fiber can also be calculated. Greens, vegetables and insects are typically 2%, 2% and 18% fat, respectively. They are typically 25%, 25% and 15% fiber, respectively. .125x2% + .042x2% + .833x18%= 15.3% fat (goal 3%-6% omnivores, much higher for carnivores) .125x25% + .042x25% + .833x15%= 16.7% fiber (goal 20%-35%, lower for carnivores) |